Hi NZ1,
I think the sharpness and the colour are wonderful.
You need to add a fill flash and maybe even a better beamer. I have found in my situation that the distances you have to shoot from and the fact that the animals tend to congregate in shady areas, I have gotten a lotbetter results whn I always use a flash. Even if your past the flashes ranges, it will put a catch light in the eyes.
http://www.rkjarman.com/naturepage/meerkat_std.htm
Here is an example where I was on the edge of the flash range, but it added a nice catchlight and still borught out some detail in the shadows.
All that said, I really think your shots are great!